Taxacom: Extent angiosperm family in the Permian

[John Grehan]

Interesting to hear the deafening silence from all those critics of
tectonic correlation about the molecular age being extended 10's of
millions of years. Seems that what is deemed OK by one research program
(fossil calibrated molecular clock)  is not acceptable from another
(tectonic correlation). (I did get one off-list note of appreciation).

John Grehan

On Wed, Aug 31, 2022 at 7:41 PM John Grehan via Taxacom <
taxacom at lists.ku.edu> wrote:

> Panbiogeographers often get a lot of flak and ridicule for tectonic
> correlations that are much older than the oldest fossil or molecular so
> called means and confidence limits. Well, here's an interesting input. Shi
> et al 2022 "Fire-prone Rhamnaceae with South African affinities in
> Cretaceous Myanmar amber" states:
>
> "We report the discovery of two exquisitely preserved fossil flower
> species, one identical to the inflorescences of the extant crown-eudicot
> Shi et al 2022. Fire-prone Rhamnaceae with South African affinities in
> Cretaceous Myanmar amber"..."genus Phylica and the other recovered as a
> sister group to Phylica, both preserved as inclusions together with burned
> plant remains in Cretaceous amber from northern Myanmar (~99 million years
> ago). These specialized flower species, named Phylica piloburmensis sp.
> nov. and Eophylica priscastellata gen. et sp. nov., exhibit traits
> identical to those of modern taxa...110–99 Ma for Eophylica priscastellata
> and 99 Ma for Phylica piloburmensis"
>
> "The unique floral architecture preserved in Eophylica and Phylica fossils
> is identical to that seen in extant members of the genus Phylica, predating
> molecular clock estimates for the genus by at least 35 Myr"
>
> The we have He & Lamont 2022 "Ancient Rhamnaceae flowers impute an origin
> for flowering plants exceeding 250-million-years ago"
>
> "Setting the molecular clock to newly described 100-million-year-old
> flowering shoots of Phylica in Burmese amber enabled us to recalibrate the
> phylogenetic history of Rhamnaceae.We traced its origin to 260 million
> years ago (Ma) that can explain its migration within and beyond Gondwana
> since that time and implies an origin for flowering plants that stretches
> well beyond 290 Ma."
>
> "The difference in estimated dates between the most recently produced
> chronograms and that described here, using these new fossils, is profound.
> Ignoring the minor error terms in each, previous results are an order of
> magnitude lower for the origin of both Phylica and Phylicieae – the new
> chronogram adds almost 100 My to their ages, as Phylica was previously
> dated at 12–24 Ma"
>
> ". The effect on the estimated age of Rhamnaceae is even more remarkable:
> it is in the order of 150 My compared with that previously reported, 74–103
> M"
>
> ". Any past statistical concerns about large levels of uncertainty
> surrounding the means pale into insignificance compared with the effect of
> the ad hoc presence/absence of fossils used to set the molecular clock, as
> shown here."
>
> So - once again new evidence points not in the direction of 'recent'
> origins, but extant angiosperm families in at least Permo-Triassic times.
> Croizat was ridiculed for his predictions of early origins for modern
> angiosperm families and even genera, but evidently this ridicule may be
> somewhat premature.
>
> John Grehan
> --
>
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