Taxacom: Neotypification and common usage
kotatsu at fripost.org
kotatsu at fripost.org
Sun Dec 11 19:18:41 CST 2022
2022
Dear Francisco and Thomas,
Thanks a lot for both your help!
Yes, this is indeed a parasitic animal. For full disclosure, it is the
species Philopterus reguli Denny, 1842, Set 1 is Najer et al. 2020
(https://nam10.safelinks.protection.outlook.com/?url=https%3A%2F%2Fphthiraptera.myspecies.info%2Fnode%2F94995&data=05%7C01%7Ctaxacom%40lists.ku.edu%7C8d6a2f3daf1d4664d1e508dadbded682%7C3c176536afe643f5b96636feabbe3c1a%7C0%7C0%7C638064047390112861%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=IR%2Bzk6I5%2FdDltcOvRDaIDXqTShmOnZiVnQCk%2BkuvmNM%3D&reserved=0), and Set 2 is Mey 2020
(https://nam10.safelinks.protection.outlook.com/?url=https%3A%2F%2Fphthiraptera.myspecies.info%2Fnode%2F95393&data=05%7C01%7Ctaxacom%40lists.ku.edu%7C8d6a2f3daf1d4664d1e508dadbded682%7C3c176536afe643f5b96636feabbe3c1a%7C0%7C0%7C638064047390112861%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=%2FYdTtA0XcOIYTuxw7rSwuWx5%2BSV9zSd6z7GjVc0itms%3D&reserved=0). Denny's book is
available on Biodiverity Heritage Library:
https://nam10.safelinks.protection.outlook.com/?url=https%3A%2F%2Fwww.biodiversitylibrary.org%2Fbibliography%2F137104&data=05%7C01%7Ctaxacom%40lists.ku.edu%7C8d6a2f3daf1d4664d1e508dadbded682%7C3c176536afe643f5b96636feabbe3c1a%7C0%7C0%7C638064047390112861%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=ClwPR4D2BaZtDjFgMZaPSB0Ynipf6tBQPxKXjAs0jeE%3D&reserved=0.
I spent some time going through other species described by Denny
yesterday. The slides are all digitized and photos are publicly
available on the NHML homepage, and I think there may be some important
clues here, as well as in the original publication.
First, the plates are maybe decisive here. The syntype does not have
these setae nor does the illustrated specimen. All other specimens of
the louse normally living on this host have these setae, including the
neotypye erected by Mey and the type series of the new species erected
by Najer et al. This is one of the reasons that Mey argues that the
syntype should be set aside, and the neotype accepted.
Second, Denny stated that he first had only one specimen of Ph. reguli,
but then obtained others. He does not use the term "type" for anything
as far as I know, but in 1852 the British Museum obtained the types of
his collection, and list Ph. reguli there, although they don't state how
many specimens they obtained for any species. The subsequent history of
other specimens examined by Denny is unknown, but at least in some cases
he thanks correspondents for allowing him to retain the specimens for
the duration of writing this monograph, so they may have been sent back
to various collectors. This is not stated for Ph. reguli.
I had a look at other analogous cases in Denny's monograph, in which he
stated that he first obtained a single specimen, and subsequently
obtained additional specimens. In all cases I could find, the end result
is the same: they are listed in the catalogue of what the NHML obtained
in 1852, and only one specimen marked "type" is present in that
collection today. Moreover, the slides are evidently mounted by the same
person, who has a style that no one else seem to use (probably this is
Bruce Cummings). In cases where Denny stated that he had many specimens,
and never mentioned any subsequent collections, there are many slides
mounted and marked in the same way at the NHML today.
Thirdly, we know from his Preface that he sent at least some of these
plates out to a colleague in Germany, who corrected them, before the
monograph was published. I also found an advertisement in the Annals of
Natural History from 1838 where he asks for help collecting lice form
British birds and mammals.
It thus seems conceivable to me, by analogy, that in these cases Denny
only had access to a single specimen of these species when he wrote his
descriptions and prepared his copperplates. After this was done, he
obtained additional specimens (which may or may not be conspecific), but
he did not consider these types. As the NHML only obtained the types,
and as far as I can tell all types they obtained were slide-mounted,
maybe this implies that Denny did not consider these specimens types,
and instead sent them back to whoever collected them. This would imply
that the syntype is the only type specimen there ever was. I know of no
way to establish whether this scenario is correct, but it is at least
plausible, which to me suggests that the syntype cannot be dismissed out
of hand.
As to whether the syntype can be identified -- yes, possibly. From what
I can see of the photo of the slide at the NHML homepage, there is
nothing to suggest that this specimen could not be identified if someone
put some effort in. Females in this genus are often very hard to
identify, but not impossible, especially if we can assume that it is one
of the other species known from the UK (where it was collected), as
there aren't that any species there, and many of the known ones can be
excluded immediately.
So based on your clarifications, and the closer examination of Denny's
book and material, it seems to me that the syntype cannot be dismissed
as a non-type without further evidence, and as -- as far as I can tell
-- the specimen may be identifiable if examined properly, it cannot be
set aside for that reason either, without further evidence. Can I then
conclude that, at present, the neotypification is unwarranted, until
someone has examined the syntype specimen and established whether it can
at all be identified?
Cheers,
Daniel
> Dear Daniel,
> I assume you are talking about animals, not about parasitic plants.
>
> For reliable statements I would need to see the original descriptions,
> and the cited sources Set 1 and Set 2.
>
> I have not understood one decisive sentence:
>
> "...and the original illustration is poor, but shows at least one
> potentially important character (setae in a certain area) which are
> absent in the illustration *and* in the remaining specimen, but
> present in the species that normally parasitizes this host."
>
> Does the illustration (1) show the setae or does the illlustration (2)
> show reliably the absence of setae?
>
> If 2 is true, then the neotype of Set 2 may be in conflict with Art.
> 75.3.5, and thus invalid.
>
> For determining the type series it is necessary to take into account
> published and unpublished evidence (Art. 72.4.1.1).
>
> The term "prevailing usage" is indeed to poorly defined in the Code
> and the Glossary to provide a useful answer to your question.
> Moderately common consensus is that for qualifying for being a name in
> prevailing usage, a relatively high number of usages should be
> observed. This is reflected in the Cases judged by the Commission in
> the Bulletin that you observed (I agree absolutely with Thomas that
> there is no "case law" in zoological nomenclature). The Cases you
> observed in the Bulletin can only suggest that term is not thought for
> very rarely mentioned species.
>
> If this helps for the moment.
>
> Best wishes
> Francisco
> Hello Daniel,
>
> A neotype is to be designated only "when no name-bearing type specimen
> (i.e. holotype, lectotype, syntype or prior neotype) is believed to be
> extant and an author considers that a name-bearing type is necessary
> to define the nominal taxon objectively".
>
> Obviously, different authors may interpret the evidence for a given
> case differently. Also, it may be impossible to decide unambiguously
> if a given specimen was seen by an author before or after a
> description was published. Where such conflicts threaten nomenclatural
> stability, it is relevant to submit an application to the Commission
> with a suggested solution.
>
> From what you have written, my impression is that the single existing
> suspected syntype is most likely a syntype -- we can never be sure,
> but it seems that the evidence is in favour. [But as mentioned by
> Francisco, a firm decision would require seeing the original work and
> possibly even consulting additional "published and unpublished
> evidence".] However, an existing syntype would mean that the neotype
> designation is invalid.
>
> Is it possible to make taxonomic sense out of the only remaining
> syntype, i.e., do you consider this specimen to be recognizable as a
> species (whether named or not yet named)? If this specimen is in such
> a condition that its taxonomic identity cannot be determined (except
> for not being conspecific with the new species established under Set
> 1), then it is possible to ask the Commission to set aside the
> name-bearing type (i.e., the existing syntype plus all other and
> probably lost syntypes) and designate a neotype. This neotype could
> then be the same as the one designated under Set 2, which would be in
> line with Art. 75.6: Conservation of prevailing usage by a neotype. If
> the Commission endorse the application, this would give the same
> result as accepting the conditions of Set 2, but there would no longer
> be any doubts. If the Commission rejects the application, it would
> have to provide an alternative solution of how to solve the
> nomenclatural instability.
>
> /Thomas
>
> -----Original Message-----
> From: kotatsu at fripost.org <kotatsu at fripost.org>
> Sent: 10. december 2022 13:48
> To: Thomas Pape <tpape at snm.ku.dk>
> Cc: taxacom <taxacom at mailman.nhm.ku.edu>; iczn-list
> <iczn-list at afriherp.org>
> Subject: Re: Taxacom: Neotypification and common usage
>
> [You don't often get email from kotatsu at fripost.org. Learn why this is
> important at https://aka.ms/LearnAboutSenderIdentification ]
>
> Hi,
>
> Thanks a lot!
>
> Set 1 seems to have operated under the assumption that the syntype
> specimen was also the only original specimen (indicating that they
> also didn't read the original description very carefully). The
> original author stated that he first had only a single specimen (which
> may or may not be the remaining syntype), but that he has subsequently
> obtained others. There is no mention of how many specimens were
> examined in total. I don't know whether this means that it could
> reasonably be argued that there was only ever a single type specimen,
> and that the other specimens could be considered non-types.
>
> I believe it is a pretty safe bet that all other specimens in the
> original type series are lost, and we will never know if the original
> set seen by the author in 1842 were all conspecific or not. The
> original author described over a hundred species, and no material of
> any of the species he described (or the species he just recorded for
> the first time from the UK) has ever turned up anywhere else but at
> the museum in London, which claims that it bought all types from this
> collection in 1852. They were originally card-mounted but at some
> point between 1852 and 1937 they were all slide-mounted by an unknown
> person. The syntype slide is mounted and labelled in the exact same
> way as other specimens from this collection, and there is no reason
> published by either set of authors (or otherwise known to me) to
> assume that this specimen was not in the collection bought in 1852 and
> slide-mounted before 1937. It is of course possible that there was a
> mix-up during the slide-mounting (for instance), and that the "true"
> syntypes are somewhere else in the London collection, mislabelled, but
> it seems the Set 1 authors did not check for this, and Set 2 were
> never even at this museum to examine the collection.
>
> If Set 1 (or anyone else) where to designate the syntype a lectotype
> by asserting that the evidence available does not rule out that this
> was one of the original specimens, does this override the neotype
> designation? I interpret 75.8 as saying that this may be the case, but
> that only discusses cases where the syntype was assumed to be lost,
> and then is rediscovered.
>
> Cheers,
> Daniel
>
>
>
>> Daniel:
>>
>> As for your question on standards and possible limits for "common
>> usage", one of the underlying principles upon which the Code is based
>> is that "There is no "case law" in zoological nomenclature. Problems
>> in nomenclature are decided by applying the Code directly, and never
>> by reference to precedent. If the Commission is called on to make a
>> ruling on a particular case, the decision relates to that case alone".
>>
>> As for the specific issue, the authors behind Set 1 base their
>> decision on the wrong assumption that the syntype specimen is the
>> holotype. This specimen is a syntype (if it was one of the original
>> specimens seen by the author in 1842), but the fact that it is the
>> only specimen recovered does not give it any priority over the other
>> (and potentially lost) specimens in the type series. Any specimen in
>> the type series is eligible for being designated as a lectotype, but
>> considering the syntype to be a holotype does not qualify as a
>> lectotype designation for designations made after 1999. Set 1
>> therefore does not solve the central issue of a possibly mixed type
>> series.
>>
>> Set 2 rests on their arguments that what has been called a syntype
>> specimen is actually NOT part of the original set of specimens seen by
>> the author in 1842. If their arguments are accepted, their designation
>> of a neotype is valid (if fulfilling other conditions provided by the
>> Code), and the name proposed by the authors of Set 1 is a junior
>> synonym. [NB: neoparatypes are not recognized by the Code.]
>>
>> A thorough assessment rests on whether or not the single syntype is
>> actually a syntype, and whether or not this specimen is in a condition
>> that allows for a reliable taxonomic decision.
>>
>> Regards,
>> Thomas
>>
>>
>> -----Original Message-----
>> From: Taxacom <taxacom-bounces at lists.ku.edu> On Behalf Of Daniel
>> Gustafsson via Taxacom
>> Sent: 10. december 2022 05:59
>> To: taxacom at lists.ku.edu
>> Subject: Taxacom: Neotypification and common usage
>>
>> Dear colleagues,
>>
>> I'm looking at a case where a parasite species was described about 180
>> years ago (1842) from what may be a mixture of species (amount
>> unknown), of which only a single syntype specimen remains.
>> Unfortunately, this syntype specimen appears to not be the same
>> species as has been reported from this host in the intervening 180
>> years. The original description is largely useless even to get to the
>> genus level, and the original illustration is poor, but shows at least
>> one potentially important character (setae in a certain area) which
>> are absent in the illustration *and* in the remaining specimen, but
>> present in the species that normally parasitizes this host. Two
>> authors in the 1960?1970s redescribed the species poorly, and this
>> redescription is of a different species than the syntype specimen,
>> which they did not examine.
>>
>> In 2020, two sets of authors discussed this species (one set in
>> reaction to the other), and reached different conclusions:
>>
>> Set 1 -- the syntype specimen (which they erroneously call the
>> 'holotype') is conspecific with the illustration and not contradicted
>> by the original description. Therefore, the name of this species can
>> follow the specimen, which thus is assumed to be from an unknown host.
>> The species of parasite that is normally found on this host thus needs
>> a new name, which they provide, including a description and
>> illustration and a review of the previous reports. They have visited
>> many of the museums where previously reported specimens are stored,
>> and invariably these are the same as the new species, and not the same
>> as the syntype specimen.
>> No further specimens conspecific with the syntype specimen have been
>> found, but the group is very poorly known so this is not a surprise.
>> These authors do not redescribe or illustrate the syntype specimen,
>> and appear to have made no efforts trying to compare this specimen
>> with other specimens in the same collection from the same collector
>> 180 years ago. The question of mix-up of specimens is thus open.
>>
>> Set 2 -- it cannot be established beyond reasonable doubt that the
>> syntype specimen is actually part of the original set of specimens
>> seen by the author in 1842, and much of his collection is lost. The
>> collection has changed hands at least once since 1842, it is unknown
>> who slide-mounted the specimens, and at least two earlier authors
>> contradict each other concerning which specimens were included in the
>> original purchase. It is therefore possible that the rest of the
>> original type series included the species normally found on this host,
>> and that the syntype specimen is either a mix-up, a contamination, or
>> some other form of error. All other reports of parasites in this genus
>> from this host have used the 1842 name, and for the purposes of
>> stability, we should continue to use this name and treat the new name
>> established by Set 1 as a subjective junior synonym. To this end, Set
>> 2 erects a neotype and four neoparatypes, which correspond to the
>> parasite species normally found on this host, as well as the one
>> illustrated and described as a new species by Set 1. Set 2 keeps the
>> door open to applying to the Commission for an official ruling if this
>> neotypification meets well-founded resistance.
>>
>> In the ~180 years between the original description and the revision,
>> there appear to be about 25?30 publications that use the old name for
>> this species. However, all except two of these are either just
>> inclusions in lists of members of a given genus, with no further
>> information added, or regional faunal lists where identity of the
>> specimens has been assumed based on host associations. Until 2020,
>> there has been no way to identify the original species either from
>> descriptions or illustrations, and as Set 1 also do not describe the
>> syntype specimen, this is still true.
>>
>> Without going into which set of authors is correct (although I'd value
>> input on this as well), I'm wondering about the process of setting
>> aside type specimens that are suddenly found not to represent the
>> common understanding of the species name. Where, if anywhere, is the
>> limit for "common usage" the stability of which could be threatened if
>> the names are changed? I've seen various case reports and opinions in
>> the Bulletin, where often authors will list a number of publications
>> within a certain framework to show that the name is in use, but is
>> there any official standard for this?
>>
>> Further, if there is an accepted official standard, is this affected
>> by the circumstances of usage? Apart from faunal lists where identity
>> is assumed based on host associations, there is no information of any
>> kind attached to this name between 1842 and the present. The 2020
>> authors also do not attach any new information beyond that the syntype
>> is not conspecific with the species normally found on the host.
>>
>> It would seem to me that this kind of usage would "count" towards any
>> kind of standard for the number of publications a name needs to appear
>> in to be commonly accepted. Is just listing a name without making any
>> effort to establish whether you're applying the name to the correct
>> taxon sufficient to assert that the name is in common, universal
>> usage?
>> Seems to me this would be rewarding laziness.
>>
>> I'd value any input on this.
>>
>> Cheers,
>> Daniel
>>
>>
>> --
>> Dr. Daniel R. Gustafsson, Research Assistant Professor Institute of
>> Zoology Guangdong Academy of Sciences, Guangzhou, China.
>>
>> Ask me about chewing lice!
>> _______________________________________________
>> Taxacom Mailing List
>>
>> Send Taxacom mailing list submissions to: taxacom at lists.ku.edu For
>> list information; to subscribe or unsubscribe, visit:
>> https://nam10.safelinks.protection.outlook.com/?url=https%3A%2F%2Feur02.safelinks.protection.outlook.com%2F%3Furl%3Dhttps%253A%252F%252Flist&data=05%7C01%7Ctaxacom%40lists.ku.edu%7C8d6a2f3daf1d4664d1e508dadbded682%7C3c176536afe643f5b96636feabbe3c1a%7C0%7C0%7C638064047390112861%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=yV0KE4FyrvCOtPj4PVlgDCtt2RPOmMXmdUGP3jKZ34E%3D&reserved=0
>> s.ku.edu%2Flistinfo%2Ftaxacom&data=05%7C01%7Ctpape%40snm.ku.dk%7Ce
>> 979e01916404cfb5aaa08dadaacc56c%7Ca3927f91cda14696af898c9f1ceffa91%7C0
>> %7C0%7C638062733691990701%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAi
>> LCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sda
>> ta=HcsJbuYZxny6kyi7TEhAjs0t32B5ywq5qx2QRIC8jGA%3D&reserved=0
>> You can reach the person managing the list at:
>> taxacom-owner at lists.ku.edu The Taxacom email archive back to 1992 can
>> be searched at:
>> https://nam10.safelinks.protection.outlook.com/?url=https%3A%2F%2Feur02.safelinks.protection.outlook.com%2F%3Furl%3Dhttps%253A%252F%252Ftaxa&data=05%7C01%7Ctaxacom%40lists.ku.edu%7C8d6a2f3daf1d4664d1e508dadbded682%7C3c176536afe643f5b96636feabbe3c1a%7C0%7C0%7C638064047390112861%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=%2FzHZqV%2FmwQo4URSuyQGsdH5ycOWDjSaoOne46NsoYx8%3D&reserved=0
>> com.markmail.org%2F&data=05%7C01%7Ctpape%40snm.ku.dk%7Ce979e019164
>> 04cfb5aaa08dadaacc56c%7Ca3927f91cda14696af898c9f1ceffa91%7C0%7C0%7C638
>> 062733691990701%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2
>> luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=%2BfpUk
>> %2Bz6TReOcTYW4FsLD2w6HZQdVXtzgpG5DzOkZCE%3D&reserved=0
>>
>> Nurturing nuance while assailing ambiguity and admiring alliteration
>> for about 35 years, 1987-2022.
>> _______________________________________________
>> Taxacom Mailing List
>>
>> Send Taxacom mailing list submissions to: taxacom at lists.ku.edu For
>> list information; to subscribe or unsubscribe, visit:
>> https://nam10.safelinks.protection.outlook.com/?url=https%3A%2F%2Feur02.safelinks.protection.outlook.com%2F%3Furl%3Dhttps%253A%252F%252Flist&data=05%7C01%7Ctaxacom%40lists.ku.edu%7C8d6a2f3daf1d4664d1e508dadbded682%7C3c176536afe643f5b96636feabbe3c1a%7C0%7C0%7C638064047390112861%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=yV0KE4FyrvCOtPj4PVlgDCtt2RPOmMXmdUGP3jKZ34E%3D&reserved=0
>> s.ku.edu%2Flistinfo%2Ftaxacom&data=05%7C01%7Ctpape%40snm.ku.dk%7Ce
>> 979e01916404cfb5aaa08dadaacc56c%7Ca3927f91cda14696af898c9f1ceffa91%7C0
>> %7C0%7C638062733691990701%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAi
>> LCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sda
>> ta=HcsJbuYZxny6kyi7TEhAjs0t32B5ywq5qx2QRIC8jGA%3D&reserved=0
>> You can reach the person managing the list at:
>> taxacom-owner at lists.ku.edu
>> The Taxacom email archive back to 1992 can be searched at:
>> https://nam10.safelinks.protection.outlook.com/?url=https%3A%2F%2Feur02.safelinks.protection.outlook.com%2F%3Furl%3Dhttps%253A%252F%252Ftaxa&data=05%7C01%7Ctaxacom%40lists.ku.edu%7C8d6a2f3daf1d4664d1e508dadbded682%7C3c176536afe643f5b96636feabbe3c1a%7C0%7C0%7C638064047390112861%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=%2FzHZqV%2FmwQo4URSuyQGsdH5ycOWDjSaoOne46NsoYx8%3D&reserved=0
>> com.markmail.org%2F&data=05%7C01%7Ctpape%40snm.ku.dk%7Ce979e019164
>> 04cfb5aaa08dadaacc56c%7Ca3927f91cda14696af898c9f1ceffa91%7C0%7C0%7C638
>> 062733691990701%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2
>> luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=%2BfpUk
>> %2Bz6TReOcTYW4FsLD2w6HZQdVXtzgpG5DzOkZCE%3D&reserved=0
>>
>> Nurturing nuance while assailing ambiguity and admiring alliteration
>> for about 35 years, 1987-2022.
>
> --
> Dr. Daniel R. Gustafsson, Research Assistant Professor Institute of
> Zoology Guangdong Academy of Sciences, Guangzhou, China.
>
> Ask me about chewing lice!
> _______________________________________________
> Taxacom Mailing List
>
> Send Taxacom mailing list submissions to: taxacom at lists.ku.edu
> For list information; to subscribe or unsubscribe, visit:
> https://lists.ku.edu/listinfo/taxacom
> You can reach the person managing the list at:
> taxacom-owner at lists.ku.edu
> The Taxacom email archive back to 1992 can be searched at:
> https://nam10.safelinks.protection.outlook.com/?url=https%3A%2F%2Ftaxacom.markmail.org%2F&data=05%7C01%7Ctaxacom%40lists.ku.edu%7C8d6a2f3daf1d4664d1e508dadbded682%7C3c176536afe643f5b96636feabbe3c1a%7C0%7C0%7C638064047390269516%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=ERnc399wwLxsO7Zxn7zRjuPXRoRvFR0dlwTKWESK7q8%3D&reserved=0
>
> Nurturing nuance while assailing ambiguity and admiring alliteration
> for about 35 years, 1987-2022.
--
Dr. Daniel R. Gustafsson, Research Assistant Professor
Institute of Zoology Guangdong Academy of Sciences, Guangzhou, China.
Ask me about chewing lice!
More information about the Taxacom
mailing list